Genetics and Colors of the Siamese Fighting Fish, Betta splendens

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Abstract
Highly colored types were proved dominant over a light-yellow type known as Betta cambodia. The 1st group included the blue B. cyana, the green B. smaragdgreen, the red B. rubra, and a light blue type. The normal wild type, B. splendens, is mixed red and brown. All these colors are produced by combinations of black, red, and yellow chromatophores and by reflecting cells called guano-phores. The recessive factor in B. cambodia did not behave in strictly Mendelian fashion but often gave a 2 dark to 1 cambodia in place of 3 to 1.

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