The Sweetognathus complex in the Permian of China: implications for evolution and homeomorphy

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Abstract
The well-exposed and fossiliferous Permian carbonates in China have yielded Early and Late Permian species of the Sweetognathus complex that permit worldwide stratigraphic evaluation of members of the group. The sporadic appearance of species of Sweetognathus and related genera throughout the Permian in western North America and Iran, in particular, may represent iterative evolution and homeomorphy. The pectiniform element morphologies of the several species are interpreted as most important for evolutionary studies and document a partial Permian biostratigraphy in China that aids in the interpretation of less complete sequences elsewhere. A new genus, Pseudosweetognathus, and four new species, Pseudosweetognathus costatus, Sweetognathus subsymmetricus, S. paraguizhouensis, and Iranognathus nudus, are described.

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