Reconstructing Triassic vegetation of eastern Australasia: a new approach for the biostratigraphy of Gondwanaland

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Abstract
As is widely recognised, fossil florules are difficult to correlate because they appear to have been more controlled by environmental and ecological factors than uniform changes which reflect the passage of geological time. Here I present a worked example of a generally applicable approach to this problem in dating and mapping fossil plant associations. Floral associations are best described and named by the classical methods of phytosociology. The three dimensional shape of an association in rocks and its palaeoecology can then be critically assessed. For example, during Middle Triassic time the following plant associations would have been encountered on a traverse from the coast to several hundred kilometres inland from the Pacific margin of Gondwanaland: Pachydermophylletum (mangrove scrub), Linguifolietum (coastal swamp woodland), Dicroidietum odontopteroidium (floodplain forest), Phoenicopsetum (levee bank scrub), Dicroidietum odontopteroidium xylopterosum (xerophytic woodland) and Johnstonietum (mallee-like woodland). The fourth dimension of fossil plant associations (time) is best assessed from the evolution of a prominent group of plants. In conjunction with the ranges of other plant megafossils, evolutionary changes can be used to define Oppel-zones. For the Late Permian to Early Jurassic of eastern Australasia, I have used the evolution of Dicroidium and leaves of related pteridosperm plants together with other species of restricted stratigraphic range or prominent in the definition of the fossil plant associations. From this evidence four Oppel-zones can be recognised. ‘Thinnfeldia’ callipteroides Oppel-zone (Chhidruan to Smithian, 250–230 × 106 years), Dicroidium zuberi Oppel-zone (Smithian to Middle Anisian, 230–220 × 106 years), D. odontopteroides Oppel-zone (Late Anisian to Ladinian, 220–210 × 106 years) and Yabeiella Oppel-zone (Carnian to Rhaetian, 210–200 × 106 years. In a complete revision of the pteridosperm form-genera Dicroidium, Johnstonia and Xylopteris, a large number of new combinations and the following new taxa are proposed; Dicroidium gouldii sp. nov., D. odontopteroides var. argenteum var. nov., D. odontopteroides var. moltenense var. nov., D. radiatum sp. nov., D. townrovii sp. nov. and Johnstonia coriacea var. obesa var. nov.