Abstract
Thaine's observation of streaming in transcellular strands of protoplasm in sieve-tubes is considered in relation to the main body of experimental facts about translocation. A theory is proposed which explains bi-directional movement, the logarithmic profile, the measured velocities, the dependence on respiration, movement down gradients of concentration and many other observations, and to some extent explains the observed rates of dry-weight transfer. Further experiments to test the theory are suggested.