Abstract
A new genus, Archepiolus, and a new species, schmidi, belonging to the family Neopseustidae and suborder Dacnonypha are described in this paper. This genus and species occupy a very primitive position in the Neopseustidae. The venation shows the most primitive features in all the Dacnonypha; also the male genitalia show a more primitive condition than that of Neopseustis meyricki Hering. Some aspects of the male genitalia are close to Mnesarchaea paracorma Meyrick of the Mnesarchaeidae.The discovery of this moth provides new morphological evidence on the problem of the ancestors of the Dacnonypha and of all Lepidoptera.

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