Abstract
This study concerns the distribution, both of populations in a landscape and of in- dividuals in a population, and the annual cycle of two species of beetles of the forest floor, Pterostichus oblongopunctatus and Philonthus decorus. The distribution pattern ofpopu- lations of Pt. oblongopunctatus in a given landscape changes very little from year to year, and less than Ph. decorus. This comment holds also within the population. Pt. oblongopunc- tatus has a more stable pattern of favourable places within its habitat than has Ph. decorus. So individuals of the latter species have to cope with a more unpredictable environment than the former. A description of the preferred habitats is given. The habitat of Pt. oblongopunctatus is determined more strictly by a number of structural and physical properties of the environment than is the case for Ph. decorus. Although density fluctuations differ con- siderably in each population in the landscape, they run parallel in the various places within the habitat of one population.

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