A pteranodontid pterosaur from the Early Cretaceous of Peru, with comments on the relationships of Cretaceous pterosaurs

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Abstract
The first occurrence of a pterosaur fossil from the Early Cretaceous of Peru is reported. The specimen consists of a nearly complete right humerus of a pteranodontid pterosaur preserved in three dimensions, presumably in a concretion. The Pteranodontidae is rediagnosed, and included genera and materials are listed. The Nyctosauridae and Dsungaripteridae are also diagnosed on the basis of postcranial characters, and the relationships of those three taxa plus the Azhdarchidae are discussed. The species Santanadactylus spixi Wellnhofer (1985) is reassigned to the Dsungaripteridae.