Abstract
A key to the beetles of the 33 families of Heteromera (Cucujoidea) is presented for the world and a summary of about 67 imaginal characters (where known) for each family is also included. The key is intended not only for those who wish to identify beetles with heteromeroid aedeagi but also for those who want to learn the existing gaps in our knowledge on this subject and problems for future researches. Sincere efforts to distinguish or discover the primitive and derivative characters and groups for all families of beetles (Coleoptera including Strepsiptera) are urgently required.

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