The structure and composition of three basic cirrus cloud types are examined through coordinated aircraft and ground-based polarization lidar and radar measurements. The cloud systems consist of a multilayered orographic cirrus, a 6-km deep cirrostratus, and a group of fibrous cirrus bands at the tropopause. The data reveal the presence of mesoscale generating regions with horizontal dimensions ranging from ∼15 km in narrow cloud bands up to ∼100 km in cirrostratus. These generating regions appear to be composed of complexes of much smaller convective structures, presumably on the ∼1-km scale of cirrus uncinus cells, and so are termed Mesoscale Uncinus Complexes (MUC). Accumulations of ice particles within cirrus, commonly referred to as precipitation trails, are associated with generating regions at or near cloud tops, but are also created by the local production of ice crystals within embedded convective impulses. Supercooled cloud droplets large enough to be detected by aircraft probes (≳5 μm ... Abstract The structure and composition of three basic cirrus cloud types are examined through coordinated aircraft and ground-based polarization lidar and radar measurements. The cloud systems consist of a multilayered orographic cirrus, a 6-km deep cirrostratus, and a group of fibrous cirrus bands at the tropopause. The data reveal the presence of mesoscale generating regions with horizontal dimensions ranging from ∼15 km in narrow cloud bands up to ∼100 km in cirrostratus. These generating regions appear to be composed of complexes of much smaller convective structures, presumably on the ∼1-km scale of cirrus uncinus cells, and so are termed Mesoscale Uncinus Complexes (MUC). Accumulations of ice particles within cirrus, commonly referred to as precipitation trails, are associated with generating regions at or near cloud tops, but are also created by the local production of ice crystals within embedded convective impulses. Supercooled cloud droplets large enough to be detected by aircraft probes (≳5 μm ...