The primitive angiosperms in the South Pacific exhibit 3 modes of pollination. The flowers of Drimys are pollinated by a wide variety of Diptera. One species of thrips (Thysanoptera) pollinate the flowers of Belliolum. Coleoptera play an important role in the reproductive biology of Zygogynum (plus primitive moths) and Degeneria. The flowers of all genera examined are adapted for outcrossing. Coleoptera-pollinated flowers display the most intricate floral movements. Diptera and Coleoptera may have been the pollinators of the early angiosperms.