Abstract
A cytologically aberrant mutant, found in Dawson's Golden Chaff winter wheat, had white glumes that were significantly longer than normal and the plant was significantly shorter than normal. It was characterized by a telokinetic pair of chromosomes. Evidence from two other lines has shown that such chromosomes arise from fragmentation of either primary or secondary univalents.The results, together with those obtained from speltoid and fatuoid studies, indicate that domestic species and varieties of cereals differ from their prototypes, not necessarily through gene substitution, but through the addition of genes that mask the effect of primitive genes still a part of the complex or through gene substitution in a chromosome other than that carrying the primitive gene in question.

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