Arnica gracilis (Compositae), a Natural Hybrid between A. latifolia and A. cordifolia

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Abstract
Arnica gracilis is a putative natural hybrid between A. cordifolia and A. latifolia. Because it is relatively common, morphologically ecologically distinct, and an autonomous apomict, A. gracilis is recognized at the specific level. Its flavonoid profile consists of fourteen compounds and is essentially a summation of the two parental profiles. Four compounds are common to both A. gracilis and A. cordifolia, three compounds are common to both A. gracilis and A. latifolia, two compounds are unique to A. gracilis, and five compounds are common to all three species. Arnica gracilis probably originated prior to or during the Pleistocene.