Abstract
It was shown that, when 14CO2, was fed to a single leafy shoot of a rooted willow cutting, activity appeared within 12 hrs. in honeydew collected from a colony of the aphid Tuberolachnus salignus (Gmelin), feeding on the opposite side of the stem 20 cm. below the base of the leafy shoot. This was shown to be due to a tangential movement of labelled materials through the bark. This tangential movement was found only in cuttings with limited root growth. Where an active root sink was present, extensive tangential movement was absent. Using two aphid colonies on either side of the stem, determinations of the speed of tangential movement have been made. The highest value obtained was 10.6 mm./hr.

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