Secretion of 18-Hydroxydeoxycorticosterone by the Rat Adrenal Gland

Abstract
An extraction and purification procedure, similar to that originally proposed by Silber et al. (11) for the fluorimetric determination of corticosterone in rat peripheral plasma, was adapted to quantitating 18-hydroxydeoxycorticosterone in arterial and adrenal venous rat plasma with the Porter- Silber reagent. 18-Hydroxydeoxycorticosterone was found to be the only 18-hydroxylated steroid present at a sufficiently high concentration in rat plasma to react with the reagent. The method can measure 1–5 μg of 18-hydroxydeoxycorticosterone in small (0.25–2.5 ml) or large (30–60 ml) volumes of adrenal vein and arterial blood plasma, respectively. Relative intensities of chromogens of standard cortisone and 18-hydroxydeoxycor.ticosterone varied little from one experiment to another over an extended period of time. Consequently,18-hydroxydeoxy corticosterone could be quantitated in terms of a cortisone standard by using the appropriate factor. Adrenal vein blood plasma from normal stressed rats had about 25 and 30 times as much corticosterone and 18-hydroxydeoxycorticosterone, respectively, as in the corresponding arterial blood plasma. The 18- hydroxydeoxycorticosterone plasma levels were approximately one half those of the corticosterone, whereas in control hypophysectomized or hypophysectomized ACTH-injected animals the levels were approximately the same. Administration of ACTH in hypophysectomized animals led to a 5- to 12-fold increase in 18-hydroxydeoxycorticosterone and corticosterone levels in adrenal vein blood plasma. 18-Hydroxydeoxycorticosterone is a natural steroid hormone biosynthesized in vivo by the rat adrenal gland, and its production, like that of corticosterone, is at least in part under ACTH control.