Origin, Development, and Morphology of the Intrapetiolar Stipules of Ervatamia divaricata

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Abstract
The embryonic leaf primordium has 2 distinct regions[long dash]the axial component which is included in the axis, and the relatively un-differentiated free limb. The 2 opposite axial components together form a sheath completely inclosing an axial core. The sheath ultimately separates from the axis and then into its components. The wings and a narrow strip of the central region of each leaf base containing the stipular strands then diverge as a single undivided piece which develops into the intrapetiolar stipule. The up-growth of the stipule from the inner face of the leaf base is initiated by divisions of the epidermal and subepidermal layers of the adaxial surface. Provascular cells differentiate acropetally in the axial component. Three bundles make up a leaf trace. The laterals branch in the axial component, and the branches ultimately form the stipular strands. Colleters are developed from the inner face of the leaf base below the stipule. No vascular tissue or laticiferous tubes are present in them. The mode of origin and nature of the intrapetiolar or axillary stipule differ from the other types of stipules studied by the writer and does not support, for this plant, Goebel''s interpretation of axillary stipules.

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