Abstract
The origin of the pectic layer of the cell wall of Scenedesmus quadricauda can be related to the activity of the nuclear envelope. During the four-nucleate stage, dense vesicles are formed by budding off from the outer nuclear membrane. These vesicles then migrate toward the plasma membrane and their contents are released from the cytoplasm by the process of reverse pinocytosis. From the released matrix the props and the net are organized; they become the easily recognized basic structure of the pectic layer of the cell wall.