Abstract
A complex of freshwater plants known collectively as Chara globularis exists through most temperate to subarctic regions of the world. This study focuses on the extent of reproductive isolation existing between various populations that differ with respect to geographical separation and external morphological features. Widespread isolation was encountered ranging from progeny of reduced vigor, through nongerminative oospores, to the complete absence of gametic exchange. There is no evidence to support the view that charophytes are ever truly cosmopolitan in the sense that local populations throughout the world are even potentially interfertile. Most “species” or even microspecies appear to consist of complexes of partially to completely genetically isolated local populations.

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