Correlations Between Relatives When Intermediates are Fittest

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Abstract
It is shown that natural selection for intermediates affects correlations between relatives, the effects being different for different sets of relatives. For additive genes, when heterozygote has superior fitness, half-sib correlations are slightly lower than the parent-offspring correlations. A simple approximate relation between the 2 is derived, and a similarity to effects of epistasis on selection response is discussed.