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Abstract
THE major broad achievements by the luminaries of science often distract from our appreciation of their significant but narrower contributions to allied fields. One example is Linus Pauling's pioneering contributions to the molecular clock ( Zuckerkandel and Pauling 1965); this work is far overshadowed by his monumental explorations of the nature of chemical bonds and the structures of proteins ( Hager 1995). Another example, which is the focus of this article, is Herman Joseph (H. J., Joe to his friends) Muller's work in speciation genetics.