Dhanya, a South-east Asian genus of ozaenine ground beetles

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Abstract
The species of the South-east Asian genus Dhanya are reviewed, and three new species are described: mulu from Sarawak; andrewesi from India; and cylindera from Philippines. Dhanya bioculata is designated as the type-species and a lectotype is designated for this species. The variety pallida is synonymized with its species seminigra. The species cavicola Moore is transferred from Pseudozaena to Itamus. Keys to the genera of South-east Asian Ozaenini and to the species of Dhanya are provided. Morphological details of Dhanya are illustrated with scanning electron micrographs and drawings, and include information on the structure of the prothoracic pits, elytral flanges, and on microsculpture. A cladistic analysis of the species is included and the relationship of the genus to its sister group, Eustra, and to the other genera of Ozaenini is discussed. Specimens of Dhanya are found under bark of fallen logs and in rotting tree stumps in lowland and upland tropical rain forest; an association with ants or termites is suggested for at least one species.