THE RUSTS OF ERIOGONUM, CHORIZANTHE, AND OXYTHECA

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Abstract
Uromyces intricatus is broken into six varieties. Var. umbellati and var. major attack montane and lowland species of Eriogonum subgen. Eueriogonum in much of western North America, and the former probably attacks Oxytheca. Vars. intricatus (Oregon, California), parvifolii (California), nivei (interior British Columbia to Arizona), and pluriporosus (mainly Montana, Wyoming, and Colorado) attack various species or species groups in Eriogonum subgen. Oregonium, the variation being partly geographic and partly due to host specialization. Var. intricatus attacks Chorizanthe spp. and Eriogonum subgen. Ganysma in California. This variety has met and intergraded with var. pluriporosus in the past but the link is now broken. Uromyces bisbyi, a microcyclic derivative of var. parvifolii, is described on Eriogonum parvifolium in California. The implications on the evolution of Eriogonum and the problem of realistic nomenclature of rusts derived from both host and geographic isolation are discussed. A table clarifies the nomenclature of reported host plants.

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