Favositid Tabulates: Evidence for Poriferan Affinity

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Abstract
Calcitic pseudomorphs of desma-like siliceous spicules found in the calcareous skeleton of a Devonian thamnoporid support the proposal of poriferan origin of at least some favositid Tabulata. These favositids arose from a group of Ordovician lithistid demosponges that adapted to toxic calcium excess in shallow, tropical marine environments by developing calicoblasts within the pinacoderm, supplementing their primary siliceous spicular skeletons with basally secreted calcium carbonate. They are tentatively recognized as an order of the subclass Sclerospongiae (class Demospongiae).