Localization of Phenolic Compounds in Tannin-secreting Cells from Sambucus racemosa L. Shoots

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Abstract
The localization of phenolic compounds was investigated in two kinds of idioblastic tannin cells: (1) mononucleate tannin cells close to the promeristem in which they are initiated, and compared with the surrounding tissue, and (2) the very long coenocytes (i.e. almost as long as the whole internode of Sambucus) and producing large amounts of phenolics. One likely pathway for the appearance of these compounds in the central vacuole is postulated from their localization: phenolic precursors occur outside the ER cisternae and enter the interior of parts of the ER cisternae which undergo fragmentation and dilatation. Many small vacuoles so formed fuse and lead to the formation of a large central vacuole.