A phylogenetic analysis of the Neotropical riodinid butterfly generaJuditha,Lemonias,ThisbeandUraneis, with a revision ofJuditha(Lepidoptera: Riodinidae: Nymphidiini)

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Abstract
A cladistic analysis is presented for all twenty‐four species in the Neotropical riodinid butterfly generaJudithaHemming,LemoniasHübner,ThisbeHübner andUraneisBates based on sixty‐nine characters of male and female morphology and external facies, and utilizingAudre dominaBates as the outgroup. All characters are illustrated. The analysis confirms the monophyly ofJudithaandUraneis, but indicates thatLemoniasis polyphyletic andThisbeis paraphyletic with respect toUraneis, leading us to synonymizeUraneiswithThisbe(syn.n.).Judithais found to be the sister clade to trueLemonias+ (Thisbe+Uraneis). A revision ofJudithais presented which includes discussions on the taxonomy, biology and distribution of its species, and illustrations of the adults and male and female genitalia of all taxa and the early stages of an exemplar,J. caucana. Eight species are recognized inJuditha, including two,J. nazaandJ. inambari, which are described as new. The following new generic combinations are made:rubigoBates is transferred fromJudithatoPachythoneBates;agaveGodman & Salvin andleucogoniaStichel are transferred fromLemoniastoPseudonymphidiaCallaghan;ochraceaMengel,theodoraGodman andalbofasciataGodman are transferred fromAudreHemming toLemonias;fenestrellaLathy is transferred fromThisbetoSynargisHübner;hyalinaButler,ucubisHewitson andincubusHall, Lamas & Willmott are transferred fromUraneistoThisbe;andoditesCramer (=phylleusAuctt.) is transferred fromSynargistoJuditha(comb.n.).