Abstract
1. The co-ordination of the swimming movement (beat) and feeding response in Aequorea forskalea has been studied with particular attention to the pathways taken by the excitation. 2. The rapid through-conducting system which co-ordinates the beat is sharply distinguished physiologically from the radial system which co-ordinates feeding. 3. The spontaneous origin of the beat and its rapid marginal conduction are both inhibited while the radial muscle is contracting. 4. Inhibition of this type has been observed in other genera of Hydromedusae

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