Response to Comment on "The Pentacovalent Phosphorus Intermediate of a Phosphoryl Transfer Reaction"

Abstract
First, the β-PGM-phosphorane complex contains two phosphoryl groups per enzyme subunit. A putative β-PGM-magnesium trifluoride complex, on the other hand, would contain one phosphoryl group per subunit. Bradford protein and Malachite Green phosphate assays, carried out in triplicate on solutions generated from washed crystals, defined an enzyme: phosphate stiochiometry of 1: 2 within 10% error. This result is consistent with the phosphorane structure and inconsistent with the magnesium trifluoride model.
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