The antheridial spike consists of 20-100 diminutive and strikingly urn-shaped fertile leaves. These are sometimes in 1 continuous series, but their development is often interrupted by the appearance of 1 or more series of larger sterile leaves. The spike may be simple or it may fork once or several times to form a dorsiventral, fanlike spray. The spike may persist and grow for several years. At any one time it may eon- tain up to 22 living antheridia, from one of but a few cells to those holding ripe spermatozoids. Antheridium development resembles that in Plagiochila asplenoides, so far as this latter was followed by Leitgeb. The nucleus of each of the spermatozoids, of which there are 25,000 or more in each antheridium, is at first globular, then pear-shaped, and finally becomes a slender cylinder or club. It has a peripheral, granular net or sometimes a series of transverse bands of chromatin. The young chromosomes, at early prophase of the last division, show series of minute component granules, the chro-momeres, as do the chromosomes of the nuclei of the young spermatozoids organized immediately after this division. No constant difference in form or size could be discovered among the chromosomes of the same or of different spermatozoids. Comparison of mitoses in ? and ? plants gave, likewise, no evidence of the presence of sex chromosomes. The blepharoplast first becomes clearly evident as a short rod when the nucleus of the young spermatozoid begins to elongate. It could not be constantly found at the poles of the last mitotic spindle. It was never seen in earlier mitoses. No clear case of vacuolization and fragmentation of the blepharoplast could be found.