Systematics of the Calopsocidae, an Oriental and Melanesian family of Psocoptera

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Abstract
As here revised, the family Calopsocidae comprises the genera Calopsocus Hagen (seventeen species), Neurosema McLachlan (seven), Callistoptera Enderlein (one) and Nemupsocus New (one), and four newly erected genera, Calosema (one species), Cyclopsocus (five), Dendropsocus (one) and Torrepsocus (two). A key to genera is provided, and keys to species of Calopsocus, Cyclopsocus and Neurosema. Ten species of Calopsocus are described, one of Calosema, four of Cyclopsocus, one of Dendropsocus, three of Neurosema and one of Torrepsocus. The type species of Dirla, D.javana Navas, is redescribed, and Dirla is synonymized with Calopsocus. Dirla furcata New is transferred to Calopsocus, Dirla navasi New to Cyclopsocus and Dirla pulleni Smithers to Torrepsocus. Calopsocus irretitus (Navas) is synonymized with Calopsocus rizali Banks. Further descriptions are provided of Callistoptera anna Enderlein, Calopsocus guttatus Smithers, Calopsocus iridescens Banks, Cyclopsocus navasi (New), Neurosema apicalis McLachlan and Torrepsocus pulleni (Smithers). The range of Calopsocus extends from Ceylon to New Caledonia, with ten species occurring in New Guinea; the remaining genera, including the most primitive, Cyclopsocus, are confined to New Guinea, where there has been considerable radiation, except that one species of Neurosema is recorded from the Philippines.

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