This chapter shows that all extant primate hands can grip small objects regardless of their shape. There is no way of describing how primate hands work without referring to models such as ‘pliers’, ‘hooks’, ‘scoops’, and so on. The human hand is relatively primitive, i.e. it does not display the morphological specialization seen in heavy non-human primates such as the great apes, whose hands are subject to the biomechanical constraints of locomotion. It is probable that the hand played a much more important evolutionary role with its fingers and palm acting as sense organs and touch agents than with its mechanical capabilities for gripping. The hand is the tool of tools because the human brain never stops devising increasingly sophisticated tools and instruments to be manipulated by it.