Since 2003, a joint research project of the universities of Frankfurt and Tübingen (Germany) has explored the changing interrelationship of environment and culture in the forest-savanna regions of West and Central Africa. This paper provides the first archaeological and archaeobotanical results of three field seasons in the rainforest of southern Cameroun. Excavations were carried out at Bwambé Hill in the vicinity of Kribi at the Atlantic coast as well as at Akonétye, Minyin and Abang Minko’o, all located in the hinterland near Ambam. At all sites a number of pit structures, which contained mostly ceramics, were excavated. In addition, at Akonétye two graves with rich ceramic and iron offerings were unearthed. They seem to be the oldest graves with iron objects yet known in Central Africa. A large body of archaeobotanical material was retrieved from the structures excavated (charcoal fragments, charred fruits and seeds, phytolith and starch samples). Of high importance is the presence of pearl millet (Pennisetum glaucum) at Bwambé Hill and Abang Minko’o in archaeological contexts dated to about 2200 bp. Charcoal and pollen data indicate that the ancient settlements were situated in a closed rainforest which was, however, massively disturbed and partly substituted by pioneer plant formations.