Abstract
By electron-microscope cytochemistry, succinate- and NADH2-tetrazolium reductase activities have been demonstrated in spermatozoa of the pulmonate gastropods, Testacella and Helix. Enzyme reaction product is restricted to the mitochondrial compartment named the secondary helix; this compartment is derived from the matrix of typical mitochondria. ‘Mitochondrial metamorphosis’ during spermiogenesis accomplishes the segregation of dehydrogenases to this matrical compartment.