Abstract
The anatomy of Cocculinella minutissima (Smith, 1904) and Osteopelta mirabilis Marshall, 1987 is described. Both species belong to the Lepetelloidea and are closely related to Addisonia Dall, 1882, the anatomy of which has been recently described by the author. Commonly derived (symapomorphic) characters of all three genera are found especially in the genital system (hermaphrodites, tcstis and ovary separated, with separated ducts, open seminal groove to the unmodified right cephalic tentacle), and in the alimentary tract (paired oesophageal glands). Distinct similarities in genital and excretory system (large right kidney isolated; hermaphrodites, separated gonads) with the Pseudococculinidae, Pyropeltidae and Lepetellidae justify a uniting superfamily Lepetelloidea, the anatomy of which is quite different from that of the Cocculinoidea (Cocculinidae and Bathy-sciadiidae)