A Review of the Leptodactylid Frogs of the Genus Pseudopaludicola in Northern South America

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Abstract
The monophyletic genus Pseudopaludicola (synapomorphy:hypertrophied antebrachial tubercle) occurs widely in northern South America where it is represented by the monophyletic Pseudopaludicola pusilla group (synapomorphy: T-shaped terminal phalanges on toes). The P. pusilla group consists of four species: P. boliviana Parker, distributed in an Amazonian arc from Surinam to Colombia to Bolivia and Paraguay; P. ceratophyes Rivero and Serna, known only from Leticia, Colombia; P. pusilla (Ruthven), distributed in northern Colombia (and south into the valley of the Rio Magdalena) and northwestern Venezuela (trans-Andean); and a new species, [P. llaneros sp. nov.] described from the llanos of northeastern Colombia and adjacent Venezuela.