THE LIFE-HISTORY AND HABITS OF THE LARCH BARK BEETLE, IPS CEMBRAE (COLEOPTERA: SCOLYTIDAE), IN THE NORTH-EAST OF SCOTLAND

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Abstract
An investigation of the life-history and habits of Ips cembrae (Heer) has been made at the Ord wood, Cawdor Estate in Nairnshire. The species is polygamous and has a 1-year life cycle with two broods a year. The first broods result from eggs laid in May and June, producing teneral adults in August and September, and the second arise from egg-laying in late September and early October; these second broods generally overwinter under bark in the larval and pupal stages, and the adults emerge the following spring. There are three larval instars before pupation. Maturation feeding of the young beetles and regeneration feeding of the old ones take place either under the bark where broods have developed, or by attacking fresh host trees.